@John_Harshman this has to be the 3rd (if not 4th) time we’ve circled this same question. These analyses have not been done. They also are confounded. Introns overcome those confounders. An intron study might get to the bottom of this. Until it is done, we just can’t say with certainty what this evidence tells us.
Please try and remember this going forward. We really always circle back to this same conclusion. Nothing has changed here since the last time we circled. I urge you to reread our large exchange about this and find some language that we both agree too, so we can avoid rehashing it over and over: John Harshman: Bottlenecks and Trans-Species Variation.