Nothing in this article even hints at supporting the proposition that biological environments store heritable information which can be passed on without the agency of DNA. They transferred nuclei which contained DNA to enucleated egg cells, enabling their development into mature frogs. This supports my contention that DNA is the only holder of heritable information.
I am willing to change my mind, but you have to provide evidence that heritable information is stored somewhere other than DNA and that the information is transmissible without the agency of DNA?
We are discussing what obtains in present life forms and not prebiotic times. In modern organisms, DNA and RNA are the only stores of heritable information. That information is not stored in organelles or other biomolecules.