I’ve been lurking on this discussion, not sure where to jump in (or too busy to). A few observations:
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Whatever sort of randomness we are talking about, there should be a Probability Density Function (PDF, or PMF). We have some empirical knowledge of this PDF, and from what I can tell, it is a mixture distribution of different types of events. Identifying those different event types would seem to be necessary before making claims about the mixture.
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Algorithmic randomness deals with sequences and compression. Randomness is this sense doesn’t apply to individual mutations, and would seem to be difficult to apply to sequences with mutations without a very high level of knowledge of biology. Any single event (or short series of events) will appear algorithmically random even if it deterministic.
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I’m not sure where this fits in, but censoring of various sorts is often present in biological data. The events we observe are often a biased sample, as some events may be unobserved for cause. Natural selection is a good example; if a creature fails to reproduce, we may never observe the mutation* responsible for the failure.
* assuming mutations are involved for this discussion.